FILE: <bc-55.htm> Pooled References GENERAL INDEX [Navigate to MAIN MENU ]
PLANT
RESISTANCE AND BIOLOGICAL PEST CONTROL (Contacts) The IPM approach
most compatible with biological control is the development of plant
resistance (Kogan 1982). Nevertheless, incompatibilities arise when mechanisms
of resistance indiscriminately affect both pests and natural enemies, or when
natural enemies are indirectly affected through their hosts or prey.
Experimental evidence of incompatibilities is shown in tomato (Duffee &
Isman 1981, Duffey & Bloem 1986, Duffey et al. 1986). This may be
illustrated with Heliothis zea, Spodoptera exigua
and the endoparasitic wasp, Hyposoter
exiguae (Vier.). When host
larvae ingest a diet with the glycoalkaloid tomatine, the development of the
parasitoid is detrimentally affected (Duffey & Bloem 1986). Kogan et al. (1992) warn that such studies demonstrate
that depending on the mechanism of resistance, natural enemies may be
detrimentally affected; and that when exploiting such mechanisms one should
weigh the risk of reducing the natural enemy load versus the benefit of the
particular resistance trait. Obrycki (1986)
studying the impact of potato glandular trichomes on Edovum puttleri
(Grissell, an egg parasitoid of the Colorado potato beetle, drew similar
conclusions. He showed that E.
puttleri readily parasitizes
L. decemlineata eggs on Solanum
tuberosum but that the
parasitoid is entrapped in glandular trichomes of Solanum berthaultii.
On S. tuberosum, egg mortality is increased not only due to
parasitism but probably also to host feeding and superparasitism. But aphid
parasitoids that are equally affected by S.
berthaultii trichomes in the
greenhouse were not greatly affected in the field, showing that moderate
levels of trichomes and the biological control of potato aphids are not
incompatible. Therefore, it is apparent that both biochemical and physical
plant defenses are potentially detrimental to natural enemies. As behavioral
adaptations of parasitoids of insects adapted to resistant lines may occur in
nature, it would be useful to identify such adapted populations when
searching for new sources of natural enemies (Kogan et al. 1992). [For
further details, please see <bc-49.htm> ] REFERENCES: <pooled.htm> [Additional references may be found at MELVYL
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